Everything about Malvaceae totally explained
Malvaceae, or the
mallow family, is a family of
flowering plants containing over 200 genera with close to 2,300 species .
The largest genera in terms of number of species include
Hibiscus (300 species),
Sterculia (250 species),
Dombeya (225 species),
Pavonia (200 species) and
Sida (200 species).
Taxonomy
The circumscription of the Malvaceae is very controversial.
The traditional Malvaceae
sensu stricto comprises a very homogeneous and cladistically monophyletic group.
Another major circumscription, Malvaceae
sensu lato, has been more recently defined on the basis that newer techniques have shown that the commonly recognised families
Bombacaceae,
Tiliaceae, and
Sterculiaceae, which have always been considered very close to Malvaceae
s.s., are not monophyletic groups. Thus the Malvaceae have been expanded to include all of these families so to have a monophyletic group. Adopting this circumscription, Malvaceae incorporates a much larger number of genera.
This article is based on the second circumscription, as presented by the Angiosperm Phylogeny Website . The Malvaceae
s.l. (hereafter simply
Malvaceae) comprise nine subfamilies. A tentative cladogram of the family is shown below. The diamond denotes a poorly supported branching (<80%).
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It is important to point out the relationships between these subfamilies are still either poorly supported or almost completely obscure, so that the very definition of the family may change dramatically as new studies are published.
If looking for information about the traditional Malvaceae
s.s., we recommend referring to
Malvoideae, the subfamily that approximately corresponds to that group.
Description
Most species are
herbs or
shrubs but some
trees and
lianas are present.
Leaves and stems
Leaves are generally
alternate, often palmately lobed or compound and palmately veined. The margin may be entire, but when
dentate a vein ends at the tip of each tooth (
malvoid teeth). Stipules are present. The
stems contain mucous canals and often also mucous cavities. Hairs are common, and are most typically stellate.
Flowers
The flowers are commonly borne in definite or indefinite axillary
inflorescences, which are often reduced to a single flower, but may also be cauliflorous, oppositifolious or terminal. They often bear supernumerary
bracts. They can be unisexual or bisexual and are generally
actinomorphic, often associated with conspicuous
bracts, forming an
epicalyx. They generally have five valvate
sepals, most frequently basally
connate. Five imbricate
petals. The
stamens are five to numerous,
connate at least at their bases, but often forming a tube around the
pistils. The
pistils are composed of two to many
connate carpels. The
ovary is superior, with axial placentation. Capitate or lobed
stigma.
The flowers have
nectaries made of many tightly packed glandular
hairs, usually positioned on the sepals.
Fruits
Most often a loculicidal
capsule, a
schizocarp or
nut.
Uses
A number are pest species in
agriculture, including
Abutilon theophrasti and
Modiola caroliniana plus others that are garden escapes.
Cotton (4 species of
Gossypium),
kenaf (
Hibiscus cannabinus) and
okra (
Abelmoschus esculentus) are important agricultural crops.
Further Information
Get more info on 'Malvaceae'.
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